The article says that the core was made from saplings that were then split to add the central horn strip, so I think one can assume that the grain was pretty vertical. As Mike said, flat grain is typically preferable in horn bow cores for stability, so perhaps the central horn strip helped with the stability issues of having vertical grain in the core. I don't think grain orientation would have much effect on the amount of stress the wood in a horn bow core can take, if this is what you meant by being 'under more pressure,' it is more of a stability issue.